Along the edge of standing water bodies with a fluctuating water level (zone 3), Carex elata can develop to be the dominant species of a tall sedge swamp. It can be linked with a tall forb community outwards (zone 2) and with a reed swamp inwards. The typical reed swamp vegetation is dominated by a few very competitive species sending a dense net of rhizomes and roots through the saturated soil and protecting it from erosion (Table 8.4). The most widespread reed species, Phragmites australis, prefers edges of standing water bodies and invades wetlands too. In wastewater treatment wetlands, it is the most effective purifying species because its thick, aerenchymcontaining rhizomes optimise the site as a habitat for micro-organisms that aid the reduction of pollutants and nutrients (Wissing 1995) (Figure 8.16). Along streams and riversides Phragmites communities are substituted by Phalaris arundinacea. Its roots and rhizoms only penetrate a shallow soil – layer and are, therefore, not very effective at protecting river banks from erosion. Bioengineering objectives are best realised when Phragmites australis is used together with Schoenoplectus lacustris in transition zones to deep water. Rampantly spreading Carex species (such as C. acutiformis) are recommended for shallow water and wetland zones. Together with Carex elata, some other tussock species and low-growing or lesscompetitive rhizome perennials are suitable for a diverse planting design in several layers as explained above.
Menyanthes trifoliate and Potentilla palustris as ground-covers prefer meso – to oligotrophic water and mediate between zones 4, 3 and 2 as well because they develop floating mats toward zone 5. A wide range of inhabitable water depths is to be found by Calltriche palustris, Crassula recurva, Hippuris vulgaris, Hottonia palustris and Sparganium spp., being able to form very diverse bodies according to water-depth. The ‘land-type’ develops a tough,
Table 8.4. Shallow water and swamp area
|
Menyanthes trifoliata |
Cosmopolitan |
Oligo- mesotrophic |
15/30 |
V-VI |
white |
Nasturtium officinale |
Eur, As |
Meso-eutrophic |
20-80 |
V-X |
white |
Pilularia globulifera |
Eur |
Mesotrophic, acid |
5-15 |
fern plant |
|
Potentilla palustris |
Cospoplolitan |
Oligo- mesotrophic |
30-40 |
V-VI |
brown-red |
Sparganium minimum |
Eur |
Mesotrophic |
5-20 |
VI-IX |
green |
Veronica beccabunga |
Eur, W-As |
Meso-eutrophic |
25-30 |
V-VIII |
blue |
Scattered plants (up to 30/10 m2) |
|||||
Alisma lanceolatum |
Eur, W-As |
Eutrophic |
50-60 |
VII-VIII |
white |
Alisma subcordatum |
N-Am |
Eutrophic |
30-50 |
VII-VIII |
white |
Alisma plantago – aquatica |
Eur, As |
Eutrophic |
40-100 |
VII-VIII |
white |
Baldellia ranunculoides |
Eur |
Oligo-mesotr. saltresist. |
5-30 |
VII-X |
white, pink |
Juncus bulbosus |
Eur |
Oligo-mesotr., acid |
5-15 |
VII-IX |
brown |
Orontium aquaticum |
N-Am |
Meso-eutrophic |
5-10 |
V-VI |
yellow |
Sagittaria latifolia |
N-Am |
Eutrophic |
50-60 |
VI-VII |
white |
Sagittaria sagittifolia |
Eur, W-As |
Eutrophic |
40-60 |
VI-VII |
white |
Monoculture species— 30/10m2) |
only to combine with shallow ground |
-cover species mentioned above (10- |
|||
Acorus calamus |
S-, E-As |
Meso-eutrophic |
60-100 |
V-VI |
white |
Bolboschoenus maritimus |
Eur, As, E – Am |
Meso-eutrophic |
30-100 |
VI-VIII |
brown |
Cladium mariscus |
Cosmopolitan |
Oligot-mesot, hard |
80-200 |
VII-XI |
brown |
Cyperus longus |
C-Eur, As |
Mesotrophic |
100-130 |
VII-IX |
brown |
Dulichium arundinaceum |
N-Am |
Mesotrophic? |
40-80 |
VII-IX |
brown |
Phragmites australis |
Cosmopolitan |
Meso-eutrophic |
100-400 |
IX-I |
brown |
Ranunculus lingua |
Eur, W-As |
Meso-eutrophic |
70/100 |
VI-VII |
yellow |
Schoenoplectus lacustris |
Eur, As |
Eutrophic |
200-250 |
VI-VIII |
brown |
Schoenopl. tabernaemontani |
Eur, As, NAm |
Eutrophic, saltresistant |
50-150 |
VI-VIII |
brown |
Sparganium erectum |
Eur, As |
Meso-eutrophic |
50-120 |
VI-VII |
green |
Sparganium simplex |
Eur, As, NAm |
Meso-eutrophic |
50-100 |
VI-VII |
green |
Typha angustifolia |
Eur, As, NAm |
Meso-eutrophic |
150-200 |
VI-X |
brown |
Typha latifolia |
Cosmopolitan |
Meso-eutrophic |
150-200 |
VI-VII |
black |
Typha laxmannii |
Eur, As |
Meso-eutrophic |
120-160 |
VI-X |
brown |
Typha minima |
Eur, As |
Oligo-mesotr., hard |
40-60 |
V-VII |
brown |
Typha shuttleworthii |
Eur |
Meso-eutrophic |
90-120 |
VI-VII |
grey-black |
Zizania latifolia |
As |
Meso-eutrophic |
100-150 |
flowering only climate |
in warm |
* Eur=Europe; As=Asia; Am=America; N=North; E=East; S=South; W=West; C=Central + Hardy if planted 50cm below water level
|
Nymphaea tuberosa |
N-Am |
Meso-eutrophic |
70-100 |
VI-X |
white |
Nymp. ‘Berthold’ |
Cultivar |
Meso-eutrophic |
40-60 |
VI-X |
pink |
Nymp. ‘Candidissima’ |
Cultivar |
Meso-eutrophic |
60-90 |
VI-X |
white |
Nymp. ‘Gladstoniana’ |
Cultivar |
Meso-eutrophic |
90-120 |
VI-X |
white |
Nymp. ‘Marliacea Carnea’ |
Cultivar |
Meso-eutrophic |
60-90 |
VI-X |
white-pink |
Nymp. ‘Moorei’ |
Cultivar |
Meso-eutrophic |
40-70 |
VI-X |
yellow |
Ranunculus aquatilis |
Cosmopolitan Mesotrophic |
50-90 |
V-IX |
white |
|
Trapa natans |
Eur, As |
Eutrophic |
50-120 |
VI-VIII |
white |
Monoculture rooted |
species—very competitive and not to combine with other species |
||||
Nymphoides peltata |
Eur, As |
Meso-eutrophic |
40-100 |
VI-VIII |
yellow |
Persicaria amphibia |
Cosmopolitan |
Meso-eutrophic |
10-100 |
VI-VIII |
pink |
Potamogeton natans |
Cosmopolitan |
Meso-eutrophic |
40-100 |
inconspicuous |
flowers |
* Eur=Europe; As=Asia; Am=America; N=North; E=East; S=South; W=West; C=Central + Requires slight proctection in very cold climates
Table 8.6. Submerged zone community
|
Myriophyllum spicatum |
Cosmopolitan |
Oligo-eutrophic, hard |
>30 inconspicuous |
flowers |
Myriophyllum verticillatum |
Cosmopolitan |
Meso-eutrophic |
>30 inconspicuous |
flowers |
Nitella flexilis |
Eur, As, Am |
Oligotrophic, acid |
>20 alga—plant |
|
Potamogeton crispus |
Cosmopolitan |
Mesotrophic |
>50 inconspicuous |
flowers |
Potamogeton lucens |
Eur, As |
Eutrophic, hard |
>50 inconspicuous |
flowers |
Floating species (without roots) |
||||
Ceratophyllum demersum |
Cosmopolian |
Eutrophic |
>40 inconspicuous |
flowers |
Ceratophyllum submersum |
Eur, As |
Eutrophic |
>40 inconspicuous |
flowers |
Utricularia vulgaris |
Cosmopolitan |
Mesotrophic, acid |
>40 VII-VIII |
yellow |
* Eur=Europe; As=Asia; Am=America; N=North; E=East; S=South; W=West; C=Central |
upright body, while the ‘submerged-type’ produces smooth, feathery leaves on soft branches, increasing the gas-exchanging plant surface. Floating branches develop to more or less dense growing blankets covering the water surface.